What is this?

I don't know what specifically you're referring to. Depending on different models of laptops some areas of the board are used and others are left blank.
 
Can't really tell by the size but it kinda looks like either a second hard drive bay that needs an adapter or possibly a slot for wireless or bluetooth? But really can't tell by the picture, not good enough.
 
It looks like mPCIe to me. Per the manual it's just "Reserved Mini-card Connector".
 
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